What Is Beekeeping?
Beekeeping is the
maintenance of bee colonies, commonly in man-made hives, by humans. A beekeeper
keeps bees to collect their honey and other products that the hive produce
(beeswax, propolis, flower pollen, bee pollen, and royal jelly), to pollinate
crops, or produce bees for sale to other beekeepers. A location where bees are
kept is called \"bee yard\".
History
Depictions of humans
collecting honey from wild bees date to 10,000 years ago. Beekeeping in pottery
vessels began about 9,000 years ago in North Africa. The domestication of bees
is shown in Egyptian art from around 4,500 years ago. Simple hives and smoke
were used and honey was stored in jars, some of which were found in the tombs
of pharaohs such as Tutankhamen. It wasn\'t until the 18th century that European
understanding of the colonies and biology of bees allowed the construction of
the moveable comb hive so that honey could be harvested without destroying the
entire colony. Traces of beeswax are found in pot shreds throughout the Middle
East beginning about 7000 BCE. In ancient Greece, aspects of the lives of bees
and beekeeping are discussed at length by Aristotle. Beekeeping was also
documented by great Roman writers; Virgil, Gaius Julius Hyginus, Varro, and
Columella. It’s been also been practiced in ancient China since antiquity.
Written artifacts by Fan Li (or Tao Zhu Gong) describes the art of beekeeping,
stressing the importance of the quality of the wooden box used and how this can
affect the quality of the honey. The Chinese word for honey (? mì,
reconstructed Old Chinese pronunciation *mjit) was borrowed from Indo-European
proto-Tocharian language, the source of \"honey\", from proto-Tocharian
*??t(?) (where *? is palatalized; cf. Tocharian B mit), cognate with English
mead.
Natural beekeeping
The natural beekeeping
movement believes that beehives are weakened by modern beekeeping and
agricultural practices, such as crop spraying, hive movement, frequent hive
inspections, artificial insemination of queens, routine medication, and sugar
water feeding. Practitioners of \"natural beekeeping\" tend to use
variations of the top-bar hive, which is a simple design that retains the
concept of having a movable comb without the use of frames or a foundation. The
horizontal top-bar hive, as championed by Marty Hardison, Michael Bush, Philip
Chandler, Dennis Murrell, and others, can be seen as a modernization of hollow
log hives, with the addition of wooden bars of specific width from which bees
hang their combs
Urban beekeeping
Related to natural
beekeeping, urban beekeeping is an attempt to revert to a less industrialized
way of obtaining honey by utilizing small-scale colonies that pollinate urban
gardens. Some have found that \"city bees\" are healthier than
\"rural bees\" because there are fewer pesticides and greater
biodiversity in the urban gardens. Also, homeowners can use their landscapes to
help feed local bee populations by planting flowers that provide nectar and
pollen. An environment of year-round, uninterrupted bloom creates an ideal
environment for colony reproduction.
Indoor beekeeping
Modern beekeepers have
experimented with raising bees indoors, in a controlled environment or indoor
observation hives. This may be done for reasons of space and monitoring, or in
the off-season. In the off-season, large commercial beekeepers may move
colonies to \"wintering\" warehouses, with fixed temperature, light and
humidity. This helps the bees remain healthy, but relatively dormant. These
relatively dormant or \"wintered\" bees survive on stored honey, and
new bees are not born.
Bee Colonies
Generally, a bee
colony consists of three types of bee:
A queen bee, which is normally the only breeding female
in the colony;
A large number of female worker bees, typically
30,000-50,000 in number;
Several male drones, ranging from thousands in a strong
hive in spring to very few during dearth or cold season.
The queen is the only
sexually mature female in the hive and all of the female worker bees and male
drones are her offspring. The queen may live for up to three years or more and
may be capable of laying half a million eggs or more in her lifetime. At the
peak of the breeding season, queen bee may be capable of laying 3,000 eggs in
one day. This would be exceptional, however; a prolific queen might peak at
2,000 eggs a day, but a more average queen might lay just 1,500 eggs per day.
The queen is fed a larger amount of royal jelly than a normal worker bee,
resulting in radically different growth and metamorphosis. The queen influences
the colony by the production and dissemination of a variety of pheromones or
\"queen substances\".
Mating of Queens
The queen emerges from
her cell after 15 days of development and she remains in the hive for 3-7 days
before venturing out on a mating flight. Subsequent mating flights may vary from
5 minutes to 30 minutes, and she may mate with several male drones on each
flight. Over several matings, possibly a dozen or more, the queen receives and
stores enough sperm from a succession of drones to fertilize hundreds of
thousands of eggs. If she does not manage to leave the hive to mate””possibly
due to bad weather or being trapped in part of the hive””she remains infertile
and becomes a drone layer, incapable of producing female worker bees. Worker
bees sometimes kill a non-performing queen and produce another. Mating takes
place at some distance from the hive and often several hundred feet in the air;
it is thought that this separates the strongest drones from the weaker ones,
ensuring that only the fastest and strongest drones get to pass on their genes.
Worker Bees
Most of the bees in a
hive are female worker bees. In summer when activity in the hive is frantic and
work goes on non-stop, the life of a worker bee may be as short as 6 weeks; in
autumn, when no brood is being raised and no nectar is being harvested, a young
bee may live for 16 weeks. Throughout their lives, worker bees\' duties are
dictated by age. For the first few weeks of their lifespan, they perform basic
chores within the hive: cleaning empty brood cells, removing debris and other
housekeeping tasks, making wax for building or repairing the comb, and feeding
larvae. Later, they may ventilate the hive or guard the entrance. Older workers
leave the hive daily, weather permitting, to forage for nectar, pollen, water,
and propolis.
Drones
Drones are the largest
bees in the hive (except for the queen), at almost twice the size of a worker
bee. They have much larger eyes than the workers have, presumably to better
locate the queen during the mating flight. They do not forage for pollen or
nectar, are unable to sting, and have no other known function than to mate with
new queens and fertilize them on their mating flights. A bee colony generally
starts to raise drones a few weeks before building queen cells so they can
supersede a failing queen or prepare for swarming. When queen-raising for the
season is over, bees in colder climates drive drones out of the hive to die,
biting and tearing their legs and wings.
Structure of a bee
colony
A domesticated bee
colony is normally housed in a rectangular hive body, within which eight to ten
parallel frames house the vertical plates of honeycomb that contain the eggs,
larvae, pupae, and food for the colony. The two outside combs at each side of
the hive tend to be exclusively used for long-term storage of honey and pollen.
Within the central
brood nest, a single frame of comb typically has a central disk of eggs, larvae
and sealed brood cells that may extend almost to the edges of the frame.
Immediately above the brood patch, an arch of pollen-filled cells extends from
side to side, and above that again a broader arch of honey-filled cells extends
to the frame tops. The pollen is protein-rich food for developing larvae, while
honey is also food but largely energy-rich rather than protein-rich. The nurse
bees that care for the developing brood secrete a special food called
\"royal jelly\" after feeding themselves on honey and pollen. The
amount of royal jelly fed to a larva determines whether it develops into a
worker bee or a queen.
The annual cycle of a bee
colony
The development of a
bee colony follows an annual cycle of growth that begins in spring with a rapid
expansion of the brood nest, as soon as pollen is available for feeding larvae.
Some production of brood may begin as early as January, even in cold winter,
but breeding accelerates towards a peak in May, producing an abundance of
harvesting bees synchronized to the main nectar flow in that region. Each race
of bees times this build-up slightly differently, depending on how the flora of
its original region blooms. Some regions of Europe have two nectar flows one in
late spring and another in late August. Other regions have only a single nectar
flow. The skill of the beekeeper lies in predicting when the nectar flow will
occur in his area and in trying to ensure that his colonies achieve a maximum
population of harvesters at exactly the right time.
Colony Reproduction:
Swarming and Supersedure
All colonies are
dependent on their queen, who is the only egg-layer. However, even the best
queens live only a few years and one or two years longevity is the norm. She
can choose whether or not to fertilize an egg as she lays it; if she does so,
it develops into a female worker bee; if she lays an unfertilized egg it
becomes a male drone. She decides which type of egg to lay depending on the
size of the open brood cell she encounters on the comb. In a small worker cell,
she lays a fertilized egg; if she finds a larger drone cell, she lays an
unfertilized drone egg.
All the time that the
queen is fertile and laying eggs she produces a variety of pheromones, which
control the behavior of the bees in the hive. These are commonly called queen
substance, but there are various pheromones with different functions. As the
queen ages, she begins to run out of stored sperm, and her pheromones begin to
fail. Inevitably, the queen begins to falter, and the bees decide to replace
her by creating a new queen from one of her worker eggs. They may do this
because she has been damaged (lost a leg or an antenna) because she has run out
of sperm and cannot lay fertilized eggs (has become a \"drone laying
queen\"), or because her pheromones have dwindled to where they cannot
control all the bees in the hive.
At this juncture, the
bees produce one or more queen cells by modifying existing worker cells that
contain a normal female egg. They then pursue one of two ways to replace the
queen: supersedure, replacing or superseding the queen without swarming, or
swarm cell production, dividing the hive into two colonies through swarming.
Factors that trigger
swarming
Some beekeepers may
monitor their colonies carefully in spring and watch for the appearance of
queen cells, which are a dramatic signal that the colony is determined to
swarm. This swarm looks for shelter. A beekeeper may capture it and introduce
it into a new hive, helping meet this need. Otherwise, it returns to a feral
state, in which case it finds shelter in a hollow tree, excavation, abandoned
chimney, or even behind shutters.
A small after-swarm
has less chance of survival and may threaten the original hive\'s survival if
the number of individuals left is unsustainable. When a hive swarms despite the
beekeeper\'s preventative efforts, good management practice is to give the
reduced hive a couple of frames of open brood with eggs. This helps replenish
the hive more quickly and gives a second opportunity to raise a queen if there
is a mating failure.
When a colony
accidentally loses its queen, workers realize that the queen is absent after as
little as an hour, as her pheromones fade in the hive. Instinctively, the
workers select cells containing eggs aged less than three days and enlarge
these cells dramatically to form \"emergency queen cells\". These
appear similar to large peanut-like structures about an inch long that hang
from the center or side of the brood combs.
The developing larva
in a queen cell is fed differently from an ordinary worker-bee; in addition to
the normal honey and pollen, she receives a great deal of royal jelly, a
special food secreted by young \"nurse bees\" from the hypopharyngeal
gland. This special food dramatically alters the growth and development of the
larva so that, after metamorphosis and pupation, it emerges from the cell as a
queen bee. The queen is the only bee in a colony that has fully developed ovaries,
and she secretes a pheromone which suppresses the normal development of ovaries
in all her workers.
HIVE FRAME
A hive frame or honey
frame is a structural element in a beehive that holds the honeycomb or brood
comb within the hive enclosure or box. The hive frame is a key part of the
modern movable-comb hive. It can be removed to inspect the bees for disease or
to extract the excess honey.
History
In 1814 Petro
Prokopovych invented the world\'s first beehive which used hive frames. Early
prototypes had a large distance between frames, and the frame lay on supporting
strips of wood. As a result, the frames were cross-attached by burr comb and
propolis to the supporting strips and were difficult to remove. In
Prokopovych\'s design, the frames were placed only in the honey chamber. In the
brood chamber, the bees built the combs in freestyle.
Johann Dzierzon
described the correct distance between combs in the brood chamber as 1½ inches
from the center of one bar to the center of the next. In 1848, Dzierzon introduced
grooves into the hive\'s side walls replacing the strips of wood to hang top
bars. The grooves were 8 mm apart and met the distance requirements for a bee
space.
In May 1852, August
von Berlepsch in Germany designed a movable frame. On October 5, 1852, in the
United States, L. L. Langstroth patented a new hive with movable frames under
US patent # US9300A. Today, the Langstroth hive is the most common design.
Types of frames:
Plastic frames: They are injected-molded out of plastic
and come in various colors. They usually come with a built-in plastic
foundation molded as one piece with cells stamped to a specific size. The
colors usually are used to distinguish types of frames within a
manufacture\'s product line (example: green for frames with drone size
foundation cells).
Queen rearing frames: Specialty frames such as cell bar
frames are used to raise new queens. The queen cups are attached
vertically to bars to encourage bees to build queen cells. Once these
cells are capped, the beekeeper moves them each to a queenless colony for
adoption.
Drone Trap frames: Some beekeepers have designed frames
specifically to encourage bees to built drone brood to cut it out as part
of an Integrated Pest Management (IPM) plan in the fight against Varroa
destructor.
Wax Foundation
Wax foundation or
honeycomb base is a plate made of wax forming the base of one honeycomb. It is
used in beekeeping to give the bees a foundation on which they can build the
honeycomb. Wax foundation is considered one of the most important inventions in
modern beekeeping.
History
Wax foundation was
invented by German Johannes Mehring in 1857, a few years after Langstroth
designed and patented the Langstroth hive in 1852. Mehring\'s wax foundation had
only the bottom of the cells, and today\'s base with the foundation of the cells
was invented by US beekeeper Samuel Wagner. The Langstroth patent did not call
for foundation and let the bees build their comb.
At first, wax
foundations were made in the wax foundation press. The first presses were made
of wood, while later presses could be made of plaster, cement, and finally
metal, which are the ones used today. Wagner also invented the wax foundation
rollers, but never perfected them; the first usable rollers were made by Amos
Root and precise mechanic Alva Washburn in 1875. In 1895. Detroit inventor
Edward Weed invented rollers that can make a wax foundation in a continuous
roll.
How to use:
Wax or plastic
foundation is inserted into a wooden frame through the top and is usually
connected to the sidebars with wire. It is not used in foundationless frames or
in plastic frames where the foundation is made of plastic and is part of the
frame itself. Foundation is not usually used in top-bar applications (where no
frames are used) such as Top Bar Hives or Warre Hives except sometimes as
starter strips.
Wax foundation has
some advantages over letting bees build their comb:
It provides a guide for bees to build straight comb.
Without foundation, the beekeeper runs the risk of having comb built
outside the Hive frame when they start, preventing its easy removal for
inspection.
Foundation built comb is usually stronger in part due
to the wiring embedded in the wax. This allows for centrifuge extraction.
Foundation allows beekeepers to increase the size of
the cells on the honeycomb. By stamping bigger cells on the foundation
than what bees would naturally build, the beekeeper is guiding bees to
build bigger cells, increasing the size of worker bees as well as the
volume of the cells for honey storage.
For these reasons, the
foundation had been used extensively in commercial operations.
Recently there has
been a large movement toward foundationless beekeeping by hobbyists for various
reasons. Some of which are listed below:
Varroa: With the expansion of Varroa destructor around
the world, some believe that natural cell size helps bees combat this
pest. Cutting out drone cells is also an effective way in an Integrated
Pest Management (IPM) plan to fight varroa.
Chemicals in the wax: Most beekeepers purchase their
foundation from beekeeping suppliers. In addition to honey, beekeepers
also need pollen and nectar from their colonies. Honey contains vitamins
and other valuable nutrients for bees. The cost of honey, including
beeswax, is variable depending on how many of these components are used
and where they are purchased. The price of honey, with a wide range of
beekeepers\' items, is also variable. These suppliers manufacture these
sheets of foundation with wax purchased from various beekeeping operations
which may have used chemicals or worked near fields where chemicals were
sprayed. With an increasing awareness of pesticides and their impact on
bees as well as the organic and natural beekeeping movements, some
beekeepers are concerned with the traceability of the wax used.
Cost and/or time: The foundation needs to be
manufactured. The beekeeper can make it or purchase it and this leads to
him or her spending time and our money on foundation or equipment to
manufacture it.
Production of raw honeycomb: Since the comb will be cut
out, it is easier to not have any wires in the comb. It is also better to
not have chemicals (pesticides) in the wax if it is going to be used for
human consumption.
A-frame has to be
wired so that the wax foundation could be inserted into it The foundation is
then soldered with the wire by using a spur embedded or electric current. Also
extant are wax foundations with an embedded wire that only need to be inserted
into the frame. Wax foundations are made in various sizes, depending on the frame
they will be inserted into. If needed, a roller knife is used to cut wax
foundations.
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